Joe Paterno, who won more games than any other major-college football coach, and who became the face of Pennsylvania State University and a symbol of integrity in collegiate athletics only to be fired during the 2011 season amid a child sexual abuse scandal that reverberated throughout the nation, died Sunday in State College, Pa. He was 85.

His family announced his death in a statement released Sunday morning. The cause was lung cancer, according to Mount Nittany Medical Center, where he had been treated. Paterno’s family announced in mid-November that he had received a diagnosis of lung cancer after a visit to a physician regarding a bronchial illness a few days earlier. He lived in State College.

During his 46 years as head coach, as he paced the sideline in his thick tinted glasses, indifferent to fashion in his white athletic socks and rolled-up baggy khaki pants, Paterno seemed as much a part of the Penn State landscape as Mount Nittany, overlooking the central Pennsylvania campus known as Happy Valley....